Ultrasensitive MAPK/ERK activation in the absence of a feedback loop in xenopus oocytes

Rémy Beaujois 1 Katia Cailliau-Maggio 1 Franck Riquet 1 Christophe Russo 1,2 Benjamin Pfeuty 3 Ralf Blossey 2 Matthieu Marin 1 Arlette Lescuyer-Rousseau 1 Jean-Pierre Vilain 1 Marc Lefranc 3 Jean-François Bodart 1
1EA4479, Université Lille 1, Villeneuve d'Ascq
2USR 3078 CNRS, Interdisciplinary Research Institute, Villeneuve d'Ascq
3PhLAM, Université Lille 1, Villeneuve d'Ascq

The MAPK cascade in Xenopus oocytes exhibits an all-or-none, ultrasensitive response, which is believed to result from a positive feedback loop. We tested the hypothesis that an ultrasensitive response may be generated for MAPK response in absence of feed-back loop, through other regulation motifs, and seek for experimental conditions where MAPK could be activated in absence of feedback. Here we describe a context where 1,10‑Phenanthroline slowly, but strongly, activates MAPK while it impairs protein synthesis in a zinc-dependent manner, abolishing any feedback loop. The induced‑MAPK response was found to be strongly ultrasensitive. This behavior may reveal a regulation motif akin to a feed-forward loop acting in vivo.

The results are presented in the context of our modeling approaches of the MAPK cascade (Russo et al., 2009, Physica-A statistical mechanics and its application, 388 (24), 5070-5080) and the dynamics of molecular mechanisms orchestrating meiosis resumption in Xenopus laevis (Pfeuty et al., 2012, Plos Computational Biology, 8(1)).








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